1962: Colgate University, Hamilton, New York, A.B. (Plant Science)
1964: Ohio University, Athens, Ohio, M.S. (Plant Science)
1968: Yale University, New Haven, Connecticut, Ph.D. (Biology) 


1998-Present: Spencer T. Olin Professor of Biology, Washington University in St. Louis
2004-Present: Co-Founder and Board Member, MOgene, LC, St. Louis, MO
2010-Present: Board of Directors, Academy of Science-St. Louis, St. Louis, MO
2015-Present: Senior Partner, Exeteur, LLS, Venture Development Company, St. Louis, MO
2016-Present: Venture Partner, Lewis & Clark Ventures, St. Louis, MO        
2016-Present: Special Assistant to Provost, Corporate Engagement, WUSTL
2010-2015: Dean of Engineering & Applied Science, Washington University in St. Louis
2008–2009: Dean of the Faculty of Arts & Sciences, Washington University in St. Louis
2005-2007: Director, Division of Biology and Biomedical Sciences, Washington University
1999-2009: Director, Washington University – Monsanto Research Program
1992-1997: Chairman, Department of Biology, University of North Carolina, Chapel Hill, NC
1989-1998: John N. Couch Professor of Biology (Inaugural), University of North Carolina, Chapel Hill, NC
1986-1989: Research Manager, Mol. Biol., Central Research Dept., DuPont Co., Wilmington, DE
1984-1986: Founding Director, Center for Gene Research and Biotechnology, Oregon State University
1968-1986: Assistant, Associate, Full Professor of Botany, Oregon State University, Corvallis, OR 

Selected Honors and Awards

1970 & 1971: Carter Award, Outstanding Undergraduate Teacher of Science, Oregon State
1979: Elizabeth P. Richie Distinguished Professor Award, Oregon State University
1989: Distinguished Alumni and Achievement Awards, Ohio University
2002: Fellow, American Association for the Advancement of Science
2002: Fellow, St. Louis Academy of Science
2007: Fellow (Inaugural), American Society of Plant Biologists
2010: Adolph E. Gude, Jr. Award, American Society of Plant Biologists 

Selected Professional Activities

1997-2001: Advisory Committee, Biological Sciences Directorate, Natl. Sci. Foundation
1991-1994: Exec Com. & President (1992) American Society of Plant Biologists
1994-1997: Inaugural Chair, Public Affairs Committee, American Society of Plant Biologists
1989-1998: Coeditor/Assoc. Editor & Editor-in-Chief (1998-2003) The Plant Cell
1991-1998: Board of Reviewing Editors, Science
1990-1999: Sci. Adv. Board, Rockefeller Foundation, Internatl. Program Rice Biotech.
1998-2005: Board of Directors, Boyce Thompson Institute for Plant Research, Ithaca, NY
2002-2007: Non Resident Fellow, Samuel Roberts Noble Foundation, Ardmore, OK
2008-2010: Member, Board of Visitors, University of Maryland, Baltimore, MD
2009-2014: Founding Partner & Director Strategic Technology Council, Nidus, St. Louis, MO